Valentine marries at the end of the 19th century, and love passes through her family from generation to generation.

Flighty Emily “Jacks” Jackson works for the British edition of Vogue magazine. Rather than pursue a relationship, Jacks regularly hooks up with her devoted ex-boyfriend, James Wildstone, and lives with Peter Simon, a gay screenwriter. When Jacks meets Argentinian photographer’s assistant Paolo Sarmiento, she assumes he is gay and tries to bring him and Peter together, unaware that Paolo is straight and in love with her.

Two tigers are separated as cubs and taken into captivity, only to be reunited years later as enemies by an explorer (Pearce) who inadvertently forces them to fight each other.