After leaving her family’s farm to study nursing in the city, a young woman finds herself on an unexpected path towards politics.

In 1911, minor stage comic, Vernon Castle meets the stage-struck Irene Foote. A few misadventures later, they marry and then abandon comedy to attempt a dancing career together. While they’re performing in Paris, an agent sees them rehearse and starts them on their brilliant career as the world’s foremost ballroom dancers. However, at the height of their fame, World War I begins.

Danny O’Neill and Hank Taylor are rival trumpeters with the Perennials, a college band, and both men are still attending college by failing their exams seven years in a row. In the midst of a performance, Danny spies Ellen Miller who ends up being made band manager. Both men compete for her affections while trying to get the other one fired.